Josef Harl | Alpine Land Snails Working Group

Mag. Josef Harl

: Email: josef.harl@nhm-wien.ac.at; Website

Natural History Museum of Vienna
Central Research Laboratories

Josef Harl performes his PhD thesis on phylogeography and morphology of Orcula spp. with special emphasis on Orula dolium. Furthermore he is performing a phylogenetic analysis of Orculidae. He contributes to many projects of the Alpine Land Snails Working Group and participated in numerous collection trips.

Publications and Congress Contributions on Alpine Land Snails

Haring, E., Kruckenhauser, L., Duda, M., Harl, J., Pokorny, P., & Sattmann, H. (2009). Tracing the change – Do phylogeographic patterns of Alpine land snails reflect past climatic changes? [Oral presentation]. 1st European BioSyst Meeting, Leiden (The Netherlands). Cite

Duda, M., Sattmann, H., Bartel, D., Haring, E., Harl, J., & Kruckenhauser, L. (2009). The genus Trochulus as an example for morphological, genetic, and ecological differentiation of Austrian Alpine Land snails. [Oral presentation]. 3. Jahrestagung NOBIS Austria, Wien (Austria). Cite

Duda, M., Kruckenhauser, L., Haring, E., Harl, J., & Sattmann, H. (2009). Genetic differentiation and shell morphology of Trochulus oreinos oreinos (Wagner 1915), Trochulus oreinos scheerpeltzi (Mikula 1957) and Trochulus hispidus (Linné 1785) in the North-Eastern Alps. [Poster]. 102^nd Conference of the German Zoological Society, Regensburg (Germany). Cite

Duda, M., Kruckenhauser, L., Haring, E., Harl, J., & Sattmann, H. (2009). Morphologic and genetic differentiation of Trochulus oreinos oreinos (Wagner 1915), Trochulus oreinos scheerpeltzi (Mikula 1957) and Trochulus hispidus (Linné 1785) in the North-Eastern Alps. [Poster]. 1st European BioSyst Meeting, Leiden (The Netherlands). Cite

Harl, J., Duda, M., Haring, E., Sattmann, H., & Kruckenhauser, L. (2009). Genetic variation in East Alpine members of the genus Orcula (Held 1837). [Poster]. 102^nd Conference of the German Zoological Society, Regensburg (Germany). Cite

Harl, J., Duda, M., Haring, E., Sattmann, H., & Kruckenhauser, L. (2009). Phylogeography of the genus Orcula (Held 1837) in the Eastern Alps. [Poster]. Annual Meeting of the GfBS, Leiden (The Netherlands). Cite

Duda, M., Kruckenhauser, L., Haring, E., Harl, J., & Sattmann, H. (2010). The snail genus Trochulus (Gastropoda: Pulmonata: Hygromiidae) as an example for morphological, genetic, and ecological differentiation of Alpine land snails. [Oral presentation]. World Congress of Malacology, Phuket (Thailand). Cite

Harl, J., Kruckenhauser, L., Sattmann, H., Duda, M., & Haring, E. (2010). Phylogeography of the East-Alpine members of the landsnail species Orcula dolium (Gastropoda: Pulmonata: Orculidae). [Poster]. NOBIS Austria Meeting, Graz (Austria). Cite

Harl, J., Kruckenhauser, L., Sattmann, H., Duda, M., & Haring, E. (2010). Phylogeography of the East-Alpine members of the landsnail species Orcula dolium (Gastropoda: Pulmonata: Orculidae. World Congress of Malacology. [Poster]. World Congress of Malacology, Phuket (Thailand). Cite

Jaksch, K., Kruckenhauser, L., Duda, M., Harl, J., Sattmann, H., & Haring, E. (2010). Phylogeography and subspecies classification of the landsnail Clausilia dubia in Eastern Austria (Gastropoda: Pulmonata: Clausiliidae). [Poster]. The Malacological Society of London, Molluscan Forum, London (UK). Cite

Zopp, L., Kruckenhauser, L., Haring, E., Harl, J., Schileyko, A., & Sattmann, H. (2010). Do geographically distinct populations of Cylindrus obtusus (Gastropoda: Pulmonata: Helicidae) differ in the mode of reproduction? [Poster]. The Malacological Society of London, Molluscan Forum, London (UK). Cite

Kruckenhauser, L., Bartel, D., Haring, E., Sattmann, H., Harl, J., & Duda, M. (2011). What is a snail species? Mitochondrial lineages of snails with a Trochulus hispidus phenotype [Talk]. Exkursionsworkshop Alpine Landschnecken, Johnsbach, Austria. Cite

Harl, J., Kruckenhauser, L., Sattmann, H., Duda, M., & Haring, E. (2011). The evolution of Orcula dolium (Gastropoda: Pulmonata: Orculidae) in the Alps and Carpathians. How the impact of the Pleistocene glaciations shaped a snail species’ distribution. [Talk]. Exkursionsworkshop Alpine Landschnecken, Johnsbach, Austria. Cite

Kirchner, S., Kruckenhauser, L., Sattmann, H., Harl, J., Duda, M., & Haring, E. (2011). Phylogeography of Pyramidula pusilla: a big mystery about a tiny snail (Gastropoda: Pulmonata: Pyramidulidae). [Poster]. 5. Jahrestagung NOBIS Austria, Salzburg (Austria), Salzburg, Austria. Cite

Kirchner, S., Kruckenhauser, L., Harl, J., Duda, M., Sattmann, H., & Haring, E. (2011). Phylogeography of Alpine populations of Pyramidula pusilla. [Talk]. Exkursionsworkshop Alpine Landschnecken, Johnsbach, Austria. Cite

Jaksch, K., Kruckenhauser, L., Duda, M., Harl, J., Sattmann, H., & Haring, E. (2011). Clausilia dubia (Draparnaud, 1805) in the Eastern Alps: phylogeography and subspecies classification (Gastropoda: Pulmonata: Clausiliidae). [Talk]. Exkursionsworkshop Alpine Landschnecken, Johnsbach, Austria. Cite

Zopp, L., Kruckenhauser, L., Haring, E., Harl, J., Schileyko, A., & Sattmann, H. (2011). Geographic differences in genital anatomy and histology among populations of Cylindrus obtusus (Draparnaud, 1805) (Gastropoda: Helicidae). [Poster]. Exkursionsworkshop Alpine Landschnecken, Johnsbach, Austria. Cite

Harl, J., Sattmann, H., & Schileyko, A. (2011). Types of the extant taxa of the landsnail genus Orcula Held 1837 (Gastropoda: Stylommatophora: Orculidae). Archiv für Molluskenkunde: International Journal of Malacology, 140(2), 175–199. https://doi.org/10.1127/arch.moll/1869-0963/140/175-199 Cite

Kruckenhauser, L., Harl, J., & Sattmann, H. (2011). Optimized drowning procedures of pulmonate landsnails allowing subsequent DNA analysis and anatomical dissections. Annalen des Naturhistorischen Museums in Wien, Serie B, 112B, 173–175. Cite

Duda, M., Sattmann, H., Haring, E., Bartel, D., Winkler, H., Harl, J., & Kruckenhauser, L. (2011). Genetic differentiation and shell morphology of Trochulus oreinos (Wagner, 1915) and T. hispidus (Linnaeus, 1758) (Pulmonata: Hygromiidae) in the northeastern Alps. Journal of Molluscan Studies, 77(1), 30–40. https://doi.org/10.1093/mollus/eyq037 Cite

Zopp, L., Kruckenhauser, L., Haring, E., Sattmann, H., Harl, J., Schileyko, A., & Sattmann, H. (2011, June). Geographic differences in genital anatomy and histology among populations of Cylindrus obtusus (Draparnaud, 1805) (Gastropoda: Helicidae). [Poster]. 6th Congress of the European Malacological Societies, Vitoria-Gasteiz (Spain). Cite

Jaksch, K., Kruckenhauser, L., Duda, M., Harl, J., Sattmann, H., & Haring, E. (2011, June). Clausilia dubia (DRAPARNAUD, 1805) in the Eastern Alps: phylogeography and subspecies classification (Gastropoda: Pulmonata: Clausiliidae). [Poster]. 6th Congress of the European Malacological Societies (CEMS), Vitoria-Gasteiz, Spain. Cite

Jaksch, K., Sattmann, H., Kruckenhauser, L., Duda, M., Harl, J., & Haring, E. (2011, November 30). Clausilia dubia (Draparnaud, 1805) in the Eastern Alps: phylogeography and the subspecies classification (Gastropoda: Pulmonata: Clausiliidae). [Poster]. The Malacological Society of London, Molluscan Forum, London (UK). Cite

Jaksch, K., Sattmann, H., Kruckenhauser, L., Duda, M., Harl, J., & Haring, E. (2011, December 2). Nomen est omen: Clausilia dubia (Gastropoda: Pulmonata: Clausiliidae) and its dubious subspecies classification. [Oral presentation]. 5. Jahrestagung NOBIS Austria, Salzburg (Austria). Cite

Kirchner, S., Kruckenhauser, L., Sattmann, H., Harl, J., Duda, M., & Haring, E. (2012, January 12). Phylogeographic analyses of Alpine populations of Pyramidula pusilla (Gastropoda: Pulmonata: Pyramidulidae) [Poster]. 6. Jahrestagung NOBIS Austria, Klagenfurt, Austria. Cite

Kirchner, S., Kruckenhauser, L., Sattmann, H., Harl, J., Duda, M., & Haring, E. (2012, January 12). Alpine populations of Pyramidula pusilla: A closer look into the species phylogeography [Poster]. 6. Jahrestagung NOBIS Austria, Klagenfurt, Austria. Cite

Jaksch, K., Sattmann, H., Kruckenhauser, L., Duda, M., Harl, J., & Haring, E. (2012, September 26). Clausilia dubia (Draparnaud, 1805) in the Eastern Alps: phylogeography and subspecies classification (Gastropoda: Pulmonata: Clausiliidae) [Poster]. Departmentssymposium Evolutionsbiologie, Univ. Vienna, Wien. Cite

Jaksch, K., Sattmann, H., Kruckenhauser, L., Duda, M., Harl, J., & Haring, E. (2012, December 1). Clausilia dubia – a (bio)diverse alpine land snail [Poster]. 6. Jahrestagung NOBIS Austria, Klagenfurt, Austria. Cite

Kirchner, S., Kruckenhauser, L., Sattmann, H., Harl, J., Duda, M., & Haring, E. (2013, February 18). Phylogeographic analysis of Alpine populations of Pyramidula pusilla. BioSyst EU 2013, Vienna, Austria. Cite

Jaksch, K., Sattmann, H., Kruckenhauser, L., Duda, M., Harl, J., & Haring, E. (2013, February 18). Biodiversity homemade⁈ - Clausilia dubia and its various subspecies. BioSyst EU 2013, Vienna, Austria. Cite

Harl, J., Páll-Gergely, B., Duda, M., Kruckenhauser, L., Sattmann, H., & Haring, E. (2013, February 18). Phylogeography of the land snail genus Orcula Held 1837 with a special focus on the Alpine species group. BioSyst EU 2013, Vienna, Austria. Cite

Cadahía-Lorenzo, L., Harl, J., Sattmann, H., Duda, M., Kruckenhauser, L., & Haring, E. (2013, February 18). New insights on the phylogenetic position of the helicid species Cylindrus obtusus based on nuclear and mitochondrial DNA marker sequences. BioSyst EU 2013, Vienna, Austria. Cite

Kruckenhauser, L., Duda, M., Bartel, D., Harl, J., Sattmann, H., & Haring, E. (2013, February 18). Divergent mitochondrial clades of snails with a Trochulus hispidus phenotype. How to confine a snail species? BioSyst EU 2013, Vienna, Austria. Cite

Jaksch, K., Sattmann, H., Kruckenhauser, L., Duda, M., Harl, J., & Haring, E. (2013, July 21). Phylogeography and subspecies classification of Clausilia dubia Draparnaud, 1805 in East Austria. World Congress of Malacology, Ponta Delgada, Azores (Portugal). Cite

Kirchner, S., Kruckenhauser, L., Sattmann, H., Harl, J., Duda, M., & Haring, E. (2013, July 21). The case of Pyramidula pusilla: adding the first pieces to the phylogeographic puzzle of Alpine populations. World Congress of Malacology, Ponta Delgada, Azores (Potugal). Cite

Cadahía-Lorenzo, L., Harl, J., Sattmann, H., Duda, M., Kruckenhauser, L., Fehér, Z., Zopp, L., & Haring, E. (2013, July 21). New insights on the phylogenetic position of the helicid species Cylindrus obtusus based on nuclear and mitochondrial DNA marker sequences. World Congress of Malacology, Ponta Delgada, Azores (Portugal). Cite

Duda, M., Kruckenhauser, L., Sattmann, H., Harl, J., & Haring, E. (2013, July 21). Different evolutionary histories in taxa: reasons and challenges A case study in Trochulus. World Congress of Malacology, Ponta Delgada, Azores (Portugal). Cite

Duda, M., Kruckenhauser, L., Bartel, D., Harl, J., Sattmann, H., & Haring, E. (2013, July 21). Divergent mitochondrial clades of snails with a Trochulus hispidus phenotype How to confine a snail species? World Congress of Malacology, Ponta Delgada, Azores (Portugal). Cite

Harl, J., Páll-Gergely, B., Duda, M., Kruckenhauser, L., Sattmann, H., & Haring, E. (2013, July 21). Phylogeography of the land snail genus Orcula Held 1837 with a special focus on the Alpine species group. World Congress of Malacology, Ponta Delgada, Azores (Portugal). Cite

Cadahía-Lorenzo, L., Harl, J., Sattmann, H., Duda, M., Kruckenhauser, L., Fehér, Z., Zopp, L., & Haring, E. (2013, November 29). The phylogenetic position of the enigmatic helicid species Cylindrus obtusus deduced from nuclear and mitochondrial DNA marker sequences. NOBIS Young Researchers´ Day, Vienna, Austria. Cite

Kirchner, S., Kruckenhauser, L., Sattmann, H., Harl, J., Duda, M., & Haring, E. (2013, November 29). Making a start on the phylogeographic puzzle of Alpine populations of Pyramidula pusilla (Gastropoda: Pulmonata: Pyramidulidae). NOBIS Young Researchers´ Day, Vienna, Austria. Cite

Cadahía, L., Harl, J., Duda, M., Sattmann, H., Kruckenhauser, L., Fehér, Z., Zopp, L., & Haring, E. (2014). New data on the phylogeny of Ariantinae (Pulmonata, Helicidae) and the systematic position of Cylindrus obtusus based on nuclear and mitochondrial DNA marker sequences. Journal of Zoological Systematics and Evolutionary Research, 52(2), 163–169. https://doi.org/10.1111/jzs.12044 Cite

Harl, J., Duda, M., Kruckenhauser, L., Sattmann, H., & Haring, E. (2014). In Search of Glacial Refuges of the Land Snail Orcula dolium (Pulmonata, Orculidae) - An Integrative Approach Using DNA Sequence and Fossil Data. PLoS ONE, 9(5), e96012. https://doi.org/10.1371/journal.pone.0096012 Cite

Kruckenhauser, L., Duda, M., Bartel, D., Sattmann, H., Harl, J., Kirchner, S., & Haring, E. (2014). Paraphyly and budding speciation in the hairy snail (Pulmonata, Hygromiidae). Zoologica Scripta, 43(3), 273–288. https://doi.org/10.1111/zsc.12046 Cite

Duda, M., Kruckenhauser, L., Sattmann, H., Harl, J., Jaksch, K., & Haring, E. (2014). Differentiation in the Trochulus hispidus complex and related taxa (Pulmonata: Hygromiidae): morphology, ecology and their relation to phylogeography. Journal of Molluscan Studies, 80(4), 371–387. https://doi.org/10.1093/mollus/eyu023 Cite

Jaksch, K., Kruckenhauser, L., Sattmann, H., Duda, M., Harl, J., & Haring, E. (2014, August 18). Biodiversity homemade?! – Clausilia dubia and its various subspecies. Workshop Alpine Land Snails 2014, Johnsbach, Austria. Cite

Kirchner, S., Kruckenhauser, L., Sattmann, H., Duda, M., Harl, J., & Haring, E. (2014, August 18). Alpine populations of Pyramidula pusilla: A closer look into the species phylogeography (Gastropoda: Pulmonata: Pyramidulidae). Workshop Alpine Land Snails 2014, Johnsbach, Austria. Cite

Cadahía, L., Harl, J., Sattmann, H., Duda, M., Luis, Kruckenhauser, L., Fehér, Z., Zopp, L., & Haring, E. (2014, August 18). Phylogenetic relationships of Cylindrus obtusus (Pulmonata, Helicidae) deduced from nuclear and mitochondrial DNA marker sequences. Workshop Alpine Land Snails 2014, Johnsbach, Austria. Cite

Harl, J., Páll-Gergely, B., Kirchner, S., Sattmann, H., Duda, M., Kruckenhauser, L., & Haring, E. (2014). Phylogeography of the land snail genus Orcula (Orculidae, Stylommatophora) with emphasis on the Eastern Alpine taxa: speciation, hybridization and morphological variation. BMC Evolutionary Biology, 14(1), 223. https://doi.org/10.1186/s12862-014-0223-y Cite

Harl, J. (2015). Phylogeography and taxonomy of the land snail genus Orcula Held, 1837 [Dissertation]. University of Vienna. Cite